|“||Godzilla and Biollante aren't monsters. It's the unscrupulous scientists who created them who are the real monsters.||„|
— Dr. Genshiro Shiragami (Godzilla vs. Biollante)
After the raid of Godzilla in 1984, a group of Japanese soldiers scoured the ruins of Tokyo in search of left behind Godzilla cells. Suddenly, the soldiers were assaulted by agents of Bio-Major, an American scientific corporation, who stole the cells. Before the agents could escape with the cells, an assassin from Saradia called SSS9 killed them and broight the samples back to Saradia. There, Dr. Shiragami was working for the Saradia Institute of Biotechnology in order to genetically engineer plants to grow in the arid deserts of the nation. This, the government hoped, would gradually take the country away from relying only on oil exports for profit, and hopefully make them the world's largest grain exporter, threatening the status of more powerful nations like the United States and Japan. Using the imported Godzilla cells, Shiragami and his botanist daughter Erika hoped to breed these immortal plants. However, before this coulf become a reality, Bio-Major bombed the laboratory, killing Erika in the process and destroying most of Shiragami's experiments. Grief-stricken, Shiragami returned to Japan, unwilling to continue his experiments.
A few years later, in 1990, Shiragami tried to keep his daughter alive by splicing what little DNA he could retrieve from her into a rosebush. He also enlisted the help of psychic Miki Saegusa to check for psychic waves coming from the rosebushes. Little did he know that both Bio-Major and SSS9 were watching from afar. When an earthquake destroyed the rosebush, Shiragami agreed to take part in a bio-weapon project with the Japanese government so that he could have access to a supply of Godzilla cells and use them to keep the roses alive. After Shiragami successfully spliced Godzilla's cells into one lone rose, it began to rapidly mutate into the monster Biollante. Two Bio-Major agents, John Lee and Michael Low, raided Shiragami's laboratory to find his notes on the bio-weapon project, the Anti-Nuclear Energy Bacteria. There, they were attacked by SSS9, but the battle was interrupted when Biollante attacked all of them, killing Low and forcing the others to retreat. Biollante then left the lab and took root in the nearby Lake Ashi, taking the form of a gigantic rose. Godzilla, freed from Mount Mihara by a Bio-Major bomb, eventually arrived at the lake, attracted by Biollante's call. Godzilla battled Biollante, seemingly killing her, much to Shiragami's dismay. However, Biollante broke apart into spores of energy and floated away, cleading Shiragami to realize she was immortal and could not die after all. Later at the Wakasa plain, Shiragami witnessed the final battle between Godzilla and Biollante in her second form. After the battle, Shiragami witnessed Erika's face appear as Biollante ascended into outer space in the form of spores, only to be shot and killed by SSS9, who was then himself killed by Sho Kuroki when he was standing on an M6000 T.C. System pad.
Doctor Shiragami appears in Godzilla: Unleashed as a member of the Global Defense Force human council. He accidentally created Biollante under General Gyozen's instructions to create an anti-monster mutagen. He also made a breakthrough in understanding what the Power-Surge crystals do to the kaiju. In the PS2 version, due to the absence of Biollante, his role is heavily reduced.
- In Godzilla: Unleashed, Shiragami's overall appearance is radically different to his film counterpart. He resembles a stereotypical American scientist with a flowing white labcoat and thick-rimmed glasses.
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