One day, Gamera's old foes reappeared, and it's up to Gamera and the JSDF to fight off the new monsters. The JSDF, anticipating heavy losses, bring in their elite units comprised of new robots and advanced ground units to help.
- All of the enemy monsters from the Gamera series appear, minus Zedus.
- This is the first Gamera video game to feature Legion and Iris as some of the game's opponents.
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