One day, Gamera's old foes reappeared, and it's up to Gamera and the JSDF to fight off the new monsters. The JSDF, anticipating heavy losses, bring in their elite units comprised of new robots and advanced ground units to help.
- Zedus is the only of Gamera's enemies to not appear in this game.
- This is the first Gamera video game to feature Legion and Iris as opponents.
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