Toto (トト is a giant turtle Toto)kaiju created by Kadokawa that first appeared in the 2006 Gamera film, Gamera the Brave. He is the reincarnation of Avant Gamera, who sacrificed himself to destroy a flock of Gyaos in 1973.
Toto's name might come from the English word "tortoise." In-universe, Toto is given his name by Toru after a nickname his late mother used to call him. Once Toto grows to adulthood, he is finally acknowledged as Gamera (ガメラ).
Toto is the reincarnation of Avant Gamera, who released his internal energy to self-destruct and destroy a flock of Gyaos in 1973. Gamera was reincarnated over a period of 33 years, eventually reappearing as a small turtle egg on top of a red gem on an island.
In 2006, 33 years after Gamera sacrificed himself to stop a flock of Gyaos, a young boy named Toru discovered a turtle egg sitting atop a red gem on an island. The boy took the egg home along with the red gem it was sitting on. The egg hatched into a baby turtle, which Toru named "Toto", a nickname his later mother used to call him. Toto surprised Toru's neighbor when he started to fly. He soon became too big for Toru to take care of in his home, so he was taken to a shed on the shoreline. He disappeared the next day, but he returned to save Toru and his friends when Zedus attacked his village. He battled Zedus and drove him off, but he was seriously injured and captured by the military. Zedus arrived in Nagoya, and was confronted by Toto after he grew larger and escaped military custody. Zedus threw Toto into a building and prepared to kill him, but Toru arrived and threw the gem found with Toto's egg into his mouth, which re-energized him and gave him the ability to fly and breathe fire. Toto battled Zedus again, and was able to tear out the beast's tongue and then destroy him with a fireball. Toto collapsed to the ground from exhaustion, and was surrounded by the military. Toru and his friends stood in front of Toto, giving him time to regain his strength and fly away. As Toto flew into the distance, Toru stared at him and said "Sayonara, Gamera."
Toto is in the facility of Amemiya throughout the manga. The researchers try to find a way to help Toto to fight Zedus.
- Main article: Toto/Gallery.
Rather than Gamera's unique roar, Toto's roar is an edited stock roar which dates back to the 1957 Universal Pictures films The Land Unknown and The Deadly Mantis and was also used for King Kong in the 1976 film.
- Baby Toto was played by a live African spurred tortoise, but when he is shown displaying varying emotions, computer generated imagery was also used.
- Toto's concept is very similar to the Godzilla characters Minilla and Godzilla Junior, both being younger members of the main monster's species. Like Godzilla Junior, Toto eventually grew up and took the place of his predecessor following his death.
This is a list of references for Toto. These citations are used to identify the reliable sources on which this article is based. These references appear inside articles in the form of superscript numbers, which look like this: 
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