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- After the Central Committee of the Aratrum approves a return to mission to Earth, the ship's passengers discover that 20,000 years have passed on the planet and that a new ecosystem has sprung up around Godzilla, now the dominant lifeform. A landing brigade headed by Colonel Eliott Leland is deployed to the former Tanzawa Forest area to carry out Haruo Sakaki's anti-Godzilla plan. They are attacked by a swarm of Servum, winged creatures which inhabit the new Godzilla-dominated ecosystem, and encounter Godzilla Filius, a subspecies of the original Godzilla that divided from his cells. Haruo enacts his plan to disable the creature's electromagnetic field and successfully kills it by overloading its body with electromagnetic energy until it explodes.. However, the original Godzilla appears, now grown to 300 meters in height, and swiftly eliminates most of the United Earth's forces. Haruo is rescued by a mysterious tribal woman named Miana. After being nursed back to health, Haruo reunites with some other survivors and is brought to the underground village of Miana's people, the Houtua, who have seemingly descended from the humans left behind on Earth 20,000 years ago. The Bilusaludo Mulu-elu Galu-gu and Rilu-elu Belu-be discover that the Houtua use Nanometal, the same metal used to construct Mechagodzilla 20,000 years ago, as a material in the construction of weapons. Miana and her twin sister Maina guide the remaining United Earth troops to the former site of Mechagodzilla's development plant at the foot of Mount Fuji, which is now the site of a vast city formed from Mechagodzilla's surviving Nanometal, which Galu-gu dubs Mechagodzilla City. Plans are put underway to lure Godzilla to Mechagodzilla City and execute him with the City's specialized anti-Godzilla Nanometal weaponry. When Godzilla resumes activity and locates the City, he is held off by Nanometal-enhanced Powered Suits called Vultures before being lured to a capture point where he is trapped by liquid Nanometal. In a scaled-up version of Haruo's previous plan, Godzilla's dorsal plates are destroyed and EMP Harpoons are injected into his body. Rather than exploding from the excess electromagnetic energy like Godzilla Filius, Godzilla converts the energy into heat. Galu-gu attempts to forcibly merge Haruo and everyone else with the Nanometal in order to let them withstand the heat long enough to destroy Godzilla, leading Haruo to the realization that Mechagodzilla City's Nanometal will assimilate the entire planet once Godzilla is destroyed. Haruo kills Galu-gu and allows Godzilla to free himself and obliterate Mechagodzilla City, although his comrade Yuko Tani is rendered brain-dead from the failed Nanometal assimilation. With the remaining Bilusaludo aboard the Aratrum calling for Haruo's head and a cult headed by the Exif archbishop Metphies rising among the surviving troops, Haruo goes into hiding with Maina. Metphies and the Exif cardinal Endurph exploit the soldiers' belief that Haruo stopping Mechagodzilla City's Nanometal was a form of divine intervention to summon forth their god, the three headed higher-dimensional being known as Ghidorah. Ghidorah destroys the Aratrum and descends to the Earth's surface to kill Godzilla. Godzilla is unable to affect Ghidorah due to the monster's extradimensional nature, and is hopelessly outmatched. With the telepathic intervention of the Houtua's goddess, Mothra, Haruo resists Metphies' influence and severs his link to Ghidorah, rendering the entity subject to this dimension's laws and allowing Godzilla to dispatch him back to his own dimension. In the aftermath, the surviving humans integrate with the Houtua, while Maina becomes pregnant with Haruo's child. When Martin Lazzari discovers a way to reactivate the Nanometal of the last surviving Vulture and potentially rebuild human civilization, Haruo understands that this along with his own vendetta against Godzilla will only renew the cycle of destruction and ruin that led to Godzilla's war with humanity and could allow Ghidorah to return to destroy the planet one day. Haruo takes Yuko's comatose Nanometal-infected body aboard the last Vulture and allows Godzilla to destroy him along with the last of the Nanometal, ensuring that humanity will find a way to survive and coexist with Godzilla as part of Houtua society. (GODZILLA: Planet of the Monsters, GODZILLA: City on the Edge of Battle, and GODZILLA: The Planet Eater)
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